The family Closteroviridae consists of diverse viruses that may pose a potential threat to the production of many agricultural crops worldwide. Members of the viruses belonging to this family cause serious diseases in economically important crops such as citrus, sugar beet, and tomato (Duffus 1973; Falk and Duffus 1988). The name Closterovirus is derived from the Greek word “Kloster” meaning filament or thread for denoting a taxonomic group of plant viruses possessing elongated, flexuous particle morphology (Bar-Joseph et al. 1979; Agranovsky 1996; Martelli et al. 2012a, b). Initially the classification was done based on the variations observed in genome size, tissue tropism, cytopathology, and relationship with vectors. The group was split into separate trichoviruses from the true closteroviruses in the early 1990s based on available genomic sequences.