This chapter shifts the focus of decentralized autonomous organizations (DAO) analysis from technical architectures to the cultural and behavioral dynamics that sustain, or undermine, decentralized governance. While smart contracts and token-based voting define formal rules, the lived reality of DAOs is shaped by narratives, rituals, identities, and informal norms. Drawing on organizational theory, anthropology, and behavioral economics, the chapter frames DAOs as socio-technical systems where code and culture co-produce governance outcomes. Key themes include the formation of collective identity through shared narratives and symbolic practices, the role of rituals and memes in fostering cohesion, and the emergence of informal hierarchies based on reputationReputation and social signaling. Behavioral dynamics such as loss aversion, herding, and polarization influence participation and deliberative quality as much as incentive design. Comparative case studies of Gitcoin, Optimism, and ENS show how cultural infrastructures interact with technical mechanisms to shape trustTrust, legitimacyLegitimacy, and governance resilience. Methodologically, the chapter advocates digital ethnography and network analysis to capture the hidden transcripts of DAO governanceDAO governancedelegation systems: informal negotiations, symbolic power, and everyday practices invisible in on-chain metrics. Ultimately, it contends that robust DAOs must be designed not only as technical protocols but also as cultural processes of meaning-making, trustTrust-building, and adaptive coordination.