<p>The in vivo circadian clock in single cyanobacteria is studied here by time-lapse fluorescence microscopy when the temperature is lowered below 25°C. We first disentangle the circadian clock behavior from the bacterial cold shock response by identifying a sequence of “death steps” based on cellular indicators. By analyzing only “alive” traces, we show that the dynamic response of individual oscillatory traces to a step-down temperature signal is described by a simple Stuart-Landau oscillator model. The same dynamical analysis applied to in vitro data (KaiC phosphorylation level following a temperature step-down) allows for extracting and comparing both clock’s responses to a temperature step down. It appears, therefore, that both oscillators go through a similar supercritical Hopf bifurcation. Finally, to quantitatively describe the temperature dependence of the resulting in vivo and in vitro Stuart-Landau parameters <InlineEquation ID="IEq1"> <InlineMediaObject> <ImageObject Color="BlackWhite" FileRef="41598_2025_97412_Article_IEq1.gif" Format="GIF" Height="19" Rendition="HTML" Resolution="72" Type="Linedraw" Width="40" /> </InlineMediaObject> <EquationSource Format="TEX">\(\mu\left(T\right)\)</EquationSource> </InlineEquation> and <InlineEquation ID="IEq2"> <InlineMediaObject> <ImageObject Color="BlackWhite" FileRef="41598_2025_97412_Article_IEq2.gif" Format="GIF" Height="19" Rendition="HTML" Resolution="72" Type="Linedraw" Width="45" /> </InlineMediaObject> <EquationSource Format="TEX">\({\omega}_{c}\left(T\right)\)</EquationSource> </InlineEquation>, we propose two simplified analytical models: temperature-dependent positive feedback or time-delayed negative feedback that is temperature compensated. Our results provide strong constraints for future models by revealing a specific time scale for transitory regimes in the cyanobacterial circadian system and its temperature dependence.</p>

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When lowering temperature, the in vivo circadian clock in cyanobacteria follows and surpasses the in vitro protein clock trough the Hopf bifurcation

  • Irina Mihalcescu,
  • Hotaka Kaji,
  • Hina Maruyama,
  • Jerôme Giraud,
  • Mathilde Van Melle-Gateau,
  • Bahram Houchmandzadeh,
  • Hiroshi Ito

摘要

The in vivo circadian clock in single cyanobacteria is studied here by time-lapse fluorescence microscopy when the temperature is lowered below 25°C. We first disentangle the circadian clock behavior from the bacterial cold shock response by identifying a sequence of “death steps” based on cellular indicators. By analyzing only “alive” traces, we show that the dynamic response of individual oscillatory traces to a step-down temperature signal is described by a simple Stuart-Landau oscillator model. The same dynamical analysis applied to in vitro data (KaiC phosphorylation level following a temperature step-down) allows for extracting and comparing both clock’s responses to a temperature step down. It appears, therefore, that both oscillators go through a similar supercritical Hopf bifurcation. Finally, to quantitatively describe the temperature dependence of the resulting in vivo and in vitro Stuart-Landau parameters \(\mu\left(T\right)\) and \({\omega}_{c}\left(T\right)\) , we propose two simplified analytical models: temperature-dependent positive feedback or time-delayed negative feedback that is temperature compensated. Our results provide strong constraints for future models by revealing a specific time scale for transitory regimes in the cyanobacterial circadian system and its temperature dependence.