Spatial heterogeneity and vertical stratification of soil nutrients in northwestern Indian forests
摘要
Soil nutrient dynamics in forest ecosystems are governed by complex interactions among vegetation cover, pedogenic gradients, and landscape-scale heterogeneity, yet depth-explicit assessments across contrasting land covers remain scarce in semi-arid alluvial systems. This study presents the first regional-scale, depth-stratified (0-30, 30-60, and 60-90 cm) assessment of twelve soil physicochemical parameters across Non-Degraded Forest and Degraded Forest land covers in the forests of Punjab and Chandigarh, northwestern India. A total of 954 soil samples from 318 locations were analysed, and linear mixed-effects modelling (LMEM) was employed to partition variance among spatial hierarchy, soil depth, and land-cover type. Soil pH ranged from 6.82 (Dasuya) to 8.76 (Ferozepur), and organic carbon from 0.27% to 1.11%, with the highest values in sub-montane northern divisions. Variance partitioning revealed that spatial hierarchy (Division/Grid random effects) accounted for the largest share of total variance (29–75%), followed by soil depth, which significantly structured organic carbon (15.23%), nitrogen (7.54%), and several micronutrients; land-cover effects were comparatively minor (0.13–2.70%). Soil depth significantly influenced pH, organic carbon (OC), nitrogen (N), potassium (K), zinc (Zn), manganese (Mn), iron (Fe), copper (Cu), and boron (B) (p < 0.001), while land cover significantly affected only pH and copper. Depth-stratified Pearson correlation analysis revealed strong pH-Fe (r = -0.42 to -0.63) and pH-Mn (r = -0.43 to -0.60) antagonisms across all profiles, confirming alkalinity-driven micronutrient immobilisation characteristic of calcareous soils. Micronutrient inter-linkages (Mn-Fe, Fe-Cu) were strongest in Non-Degraded Forest surface horizons and progressively weakened with depth, with Degraded Forest subsoils showing minimal nutrient coupling. These findings demonstrate that landscape-scale pedogenic heterogeneity and vertical biogeochemical gradients exert stronger control over nutrient distribution than categorical vegetation status alone, and underscore the need for depth-explicit soil monitoring frameworks and organic matter conservation strategies in degraded alluvial forest soils of northwestern India.