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<p>Numerous vertebrate fossil footprints have been discovered in sidewalk slabs on the campus of Islamic Azad University in Neyshabur City, northeast Iran. These slabs consist of red to brown siltstones and sandstones, with footprints preserved on bedding planes as either epireliefs or hyporeliefs. The slabs were quarried from a flagstone mine in the early Eocene deposits located east of Neyshabur. Various types of vertebrate trace fossils have been identified, including avian, perissodactyl, carnivore, and fish ichnites. Avian tracks range from small to large footprints and are assigned to <i>Avipeda phoenix</i>, <i>Gruipeda filiportatis</i>, and <i>Gruipeda fuenzalidae</i>. Tridactyl mammal footprints, <i>Moropopus elongatus</i> and <i>Plagiolophustipus montfalcoensis</i>, are attributed to Rhinocerotoidea and early Equidae, respectively. <i>Quiritipes impendens</i> is only carnivore footprint and is characterized by tetradactyl imprints. A swimming trace of fish (<i>Undichna</i> isp.) is preserved as large, sinuous, lobed ridges. In addition, other vertebrate footprints, including small pentadactyl and monodactyl imprints, are present but could not be assigned to specific ichnotaxa due to insufficient diagnostic features. Neyshabur is well known as a major tourist destination in northeastern Iran, and the fossil track-bearing sidewalks of Islamic Azad University represent a unique urban paleontological site with potential scientific, educational, and geotourism significance for tourists, travelers, academics, and students.</p>

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Early Eocene Vertebrate Tracks from Sidewalks in Neyshabur City, Northeastern Iran: An Urban Paleontological Record

  • 期刊论文

  • 发表日期：

    2026年5月6日

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  • Nasrollah Abbassi,
  • Yasaman Yasini Nasab,
  • Saeid Bakhtiari,
  • Javad Safehian

摘要

Numerous vertebrate fossil footprints have been discovered in sidewalk slabs on the campus of Islamic Azad University in Neyshabur City, northeast Iran. These slabs consist of red to brown siltstones and sandstones, with footprints preserved on bedding planes as either epireliefs or hyporeliefs. The slabs were quarried from a flagstone mine in the early Eocene deposits located east of Neyshabur. Various types of vertebrate trace fossils have been identified, including avian, perissodactyl, carnivore, and fish ichnites. Avian tracks range from small to large footprints and are assigned to Avipeda phoenix, Gruipeda filiportatis, and Gruipeda fuenzalidae. Tridactyl mammal footprints, Moropopus elongatus and Plagiolophustipus montfalcoensis, are attributed to Rhinocerotoidea and early Equidae, respectively. Quiritipes impendens is only carnivore footprint and is characterized by tetradactyl imprints. A swimming trace of fish (Undichna isp.) is preserved as large, sinuous, lobed ridges. In addition, other vertebrate footprints, including small pentadactyl and monodactyl imprints, are present but could not be assigned to specific ichnotaxa due to insufficient diagnostic features. Neyshabur is well known as a major tourist destination in northeastern Iran, and the fossil track-bearing sidewalks of Islamic Azad University represent a unique urban paleontological site with potential scientific, educational, and geotourism significance for tourists, travelers, academics, and students.